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Following the recent capture of an individual (dubbed "The Rookie"), Commissioner Gordon oversees their transfer to Stryker's Island in Metropolis. As the Rookie has developed an ability to absorb superpowers, Gordon offers the incarcerated Lex Luthor a reduced prison sentence in exchange for monitoring them and assessing their potential threat. Mercy Graves arrives and helps break Luthor and the Rookie out, alongside several other imprisoned villains, with the Justice League in pursuit. Meanwhile, the Joker and Harley Quinn steal several items from Wayne Tech, with Batman giving chase. The combined group of villains are defeated by the sudden arrival of the "Justice Syndicate", a superhero team from the parallel world of Earth-3, with only a few escaping. Before departing, Harley witnesses the Syndicate forcibly warp the Justice League away.
The Syndicate claim that the Justice League are on an off-world mission, becoming their new replacements, with leader Ultraman posing as reporter Kent Clarkson to feed false information to the public. Suspicious of the Syndicate, Luthor begins uniting a massive force of villains to expose the truth and defeat the Syndicate so that they might take over. Lois Lane and Jimmy Olsen also begin their own independent investigation. During their research, the villains discover the Justice Syndicate is actually the "Crime Syndicate", a group of Earth-3's greatest criminals, and that they are searching for something on Earth, using their superhero personas to gain easy access to classified information and locations. Once a large enough group has been assembled, the united Legion of Doom launch an attack on the Crime Syndicate on top of LexCorp, but Luthor betrays everyone, planning to use the Syndicate's technology to be rid of them all in order to rule the Earth alone.
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