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Age of Calamity is set 100 years before the events of The Legend of Zelda: Breath of the Wild, in an alternate timeline. The story depicts the Great Calamity, a great war previously alluded to in Breath of the Wild between the armies of Hyrule and the forces of Calamity Ganon that resulted in the collapse of the kingdom.
During the Great Calamity, a small Guardian awakens within Hyrule Castle. Seeing the destruction being caused, it escapes through a time portal, followed unbeknownst by a portion of Calamity Ganon's Malice. Arriving before the Calamity began, the Guardian is found by Link during battle. When it uses Impa's Sheikah Slate to raise a Sheikah Tower, they take it to Sheikah engineers Purah and Robbie for study, learning it came from a future in which Ganon reawakens. Being informed of this, King Rhoam sends Link and Zelda to meet with the four candidates selected to pilot the Divine Beasts – Mipha, Daruk, Urbosa, and Revali.
With the Champions assembled, King Rhoam commands them to find a knight able to wield the Master Sword, which rests in Korok Forest. There, they encounter a dark prophet named Astor who seeks to revive and control Calamity Ganon, but Link draws the Master Sword and forces Astor to withdraw. With Purah's research revealing the presence of other towers hidden underground, the Champions set out to activate the towers and in turn using the maps provided by said towers to lead a preemptive strike on the Yiga Clan hideout, forcing Yiga leader Master Kogha to retreat. Learning from the Guardian's records that Calamity Ganon will return on Zelda's birthday, King Rhoam sends her for training in hopes of awakening her sealing magic in time, while deploying the Divine Beasts to prepare for the Calamity's impending return.
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